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After a hive-minded alien race called the Boov conquer the Earth, they relocate the planet's human population -- all except for a little girl named Tip, who's managed to hide from the aliens. When Tip meets a fugitive Boov called Oh, there's mutual distrust. However, Oh is not like his comrades; he craves friendship and fun. As their distrust fades, the pair set out together to find Tip's mother, but, unbeknown to them, the Gorg -- enemies of the Boov -- are en route.